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Director of Studies: Professor Michael Cunliffe (micnli@mba.ac.uk), (BMA)

2nd Supervisor:  Dr George Littlejohn  

3rd Supervisor: Mr Andy Cameron,  

Applications are invited for a 3.5 years PhD studentship. The studentship will start on 1 October 2026

Scientific background
Seagrass meadows, such as those formed by Zostera marina (common eelgrass), are vital marine ecosystems that support biodiversity, stabilise sediments and sequester carbon. However, these meadows are increasingly threatened by climate change, pollution and disease. Emerging evidence highlights the role of microbial pathogens, especially the pseudofungus Phytophthora gemini (a fungus-like oomycete), in undermining restoration efforts by infecting eelgrass seeds and seedlings. This project addresses a critical knowledge gap in the biology and ecology of seagrass pathogens and their impact on restoration success, offering timely insights into disease mitigation strategies.

Research methodology
The PhD researcher will investigate the infection biology of Phytophthora gemini and its interaction with Zostera marina through a combination of laboratory assays, microscopy, and molecular techniques (e.g. qPCR, metabarcoding) to characterise pathogen life cycles, host responses and environmental tolerances. Fieldwork will be conducted at selected coastal sites to assess natural infection dynamics and collect samples for analysis. Restoration trials will be designed to test seed treatments and planting strategies under controlled and semi-natural conditions. The project will be based at the Marine Biological Association (MBA) in Â鶹´«Ã½, with additional work at the Â鶹´«Ã½ and the Ocean Conservation Trust. There will be opportunities for collaboration across UK and international research networks.

The researcher will gain interdisciplinary training in microbiology, marine ecology, plant pathology, molecular biology and restoration science, with a strong emphasis on applied ecological research. They will develop skills in experimental design, statistical analysis and scientific communication. Opportunities include attending international conferences, publishing in peer-reviewed journals and engaging with stakeholders involved in seagrass restoration. The MBA offers a vibrant research environment with access to cutting-edge facilities and mentoring from experienced scientists.

We seek an enthusiastic individual with a strong interest in marine biology, plant–microbe interactions and ecological restoration. A background in biology, environmental science or a related discipline is desirable. Prior experience in microbiology, molecular techniques, microscopy or fieldwork is advantageous but not essential. The ideal candidate will be curious, collaborative and motivated to contribute to ecological research with real-world impact.

Diver P et al & Cunliffe M (2025). Cell morphological plasticity in response to substrate availability of a cosmopolitan polymorphic yeast from the open ocean. Mycologia 117 95–109.

Chrismas N et al & Cunliffe M (2024) Metatranscriptomics reveals diversity of symbiotic interaction and mechanisms of carbon exchange in the marine cyanolichen Lichina pygmaea. New Phytologist 241 2243-2257.

Laundon D et al & Cunliffe M (2022) A cellular and molecular atlas reveals the basis of chytrid development eLife 11:e73933.

Littlejohn G et al  & Grant M (2021) Chloroplast immunity illuminated. New Phytologist 229 3088-3107.

Govers LL et al (2016) Marine Phytophthora species can hamper conservation and restoration of vegetated coastal ecosystems. Proc. R. Soc. B.28320160812

If your first language is not English, you will need to meet the minimum English requirements for the programme, IELTS Academic score of 6.5 (with no less than 5.5 in each component test area) or equivalent.  

Successful candidates who meet UKRI’s eligibility criteria will be awarded a fully-funded ARIES studentship of fees, maintenance stipend, £20,780 p.a. for 2025/26 (2026/27 rate TBC) and research costs. 

A limited number of ARIES studentships are available to International applicants. Please note however that ARIES funding does not cover additional costs associated with relocation to, and living in, the UK.

NB: The studentship is supported for 3.5 years of the four-year registration period. The subsequent 6 months of registration is a self-funded ‘writing-up’ period. 

ARIES is committed to in all areas of its operation. We encourage applications from all sections of the community regardless of gender, ethnicity, disability, age, sexual orientation, and transgender status. Projects have been developed with consideration of a safe, inclusive, and appropriate research and fieldwork environment. Academic qualifications are considered alongside non-academic experience, with equal weighting given to experience and potential. 

For further information, please visit

If you wish to discuss this project further informally, please contact Professor Michael Cunliffe; micnli@mba.ac.uk.

Please see our apply for a postgraduate research programme page for a list of supporting documents to upload with your application. 

For more information on the admissions process generally, please visit our apply for a postgraduate research programme page or contact the Doctoral College .

Shortlisted candidates will be invited for interview after the deadline. We regret that we may not be able to respond to all applications. Applicants who have not received a response within six weeks of the closing date should consider their application has been unsuccessful on this occasion.